经SEMA3A-NRP1通路探讨补髓丹调控NSCs-Exos干预PMOP作用机制
Exploration of the mechanism of Busui DAN in regulating NSCs-Exos in PMOP based on SEMA3A-NRP1 axis
  
DOI:10.3969/j.issn.1006-7108.2025.05.002
中文关键词:  绝经后骨质疏松症  补髓丹  神经干细胞来源的外泌体  SEMA3A-NRP1通路
英文关键词:neural stem cells  exosomes  Busui DAN  postmenopausal osteoporosis  SEMA3A-NRP1 axis
基金项目:国家自然科学青年基金(2004229);中国博士后科学基金面上资助“地区专项支持计划”(2021MD703844);中国博士后科学基金特别资助(站中)(2022T150285);2024年度辽宁省教育厅高校基本科研项目储备项目(2024-JYTCB-071);国家中医药管理局高水平中医药重点学科(中医基础理论)项目[国中医药人教函(2023)85号]
作者单位
唐泊文  李佳 孙丽 王静 郑洪新* 王庆谚* 辽宁中医药大学辽宁 沈阳 110847 
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中文摘要:
      目的 根据中医学“肾-骨-髓-脑”相关理论,基于SEMA3A-NRP1通路探究补髓丹调控神经干细胞来源的外泌体(NSCs-Exos)干预绝经后骨质疏松(postmenopausal osteoporosis,PMOP)的作用机制。方法 体外培养外源性大鼠神经干细胞(neural stem cell,NSCs),并使用中药复方补髓丹(BSD)干预,培育14 d后提取外泌体(NSCs-Exos),分为NSCs来源的外泌体组[(NSCs-Exos)组]和补髓丹干预NSCs的外泌体组[NSCs-Exos(BSD)组]。随后进行透射电镜(TEM)、粒径分析(NTA)和Alix、CD9、CD63外泌体标志蛋白检测;动物实验:将SD大鼠除Control组外行卵巢摘除术(OVX)造模,分为OVX组、BSD组、NSCs-Exos组、NSCs-Exos(BSD)组、NSCs-Exos+BSD组、NSCs-Exos(BSD)+BSD组。给药组进行灌胃给药,外泌体组通过尾静脉注射相应外泌体,治疗结束后取血清、肾、股骨、脑。进行骨密度、血清ALP/TRACP和HE染色检测,分析对骨密度和肾、股骨、脑结构形态的影响;运用qRT-PCR法、ELISA法、Western Blot法、WES法分析对肾、股骨、脑的SEMA3A、NRP1、Runx2、β-catenin表达量的影响;同期开展DIR小动物活体追踪实验,观察NSCs-Exos的靶向性。结果 体外实验所提取的NSCs-Exos呈现圆形颗粒,清晰可见;在CD9、CD63和Alix中均有表达,符合外泌体的特征。与NSCs-Exos组相比,NSCs-Exos(BSD)组的形态更加饱满,CD9、CD63和Alix表达量更高。动物实验:与对照组相比,OVX模型组大鼠骨小梁结构紊乱,股骨骨密度及血清ALP表达均明显降低,TRACP表达升高,SEMA3A、NRP1、Runx2、β-catenin表达明显降低(P<0.05),说明造模成功;与OVX模型组相比,治疗组股骨骨密度明显升高、骨小梁数量和之间的排列得到了改善,血清ALP表达升高,TRACP表达降低,SEMA3A、NRP1、Runx2、β-catenin表达升高(P<0.05),以NSCs-Exos(BSD)+BSD组效果最佳。并在DIR实验中发现,NSCs-Exos在肾、骨、脑组织中表达较强的荧光信号,验证了其靶向性。结论 补髓丹可以通过NSCs-Exos调控SEMA3A-NRP1通路改善骨代谢,从而防治PMOP,解析“肾-骨-髓-脑”理论相关科学内涵。
英文摘要:
      Objective Based on the theory of kidney-bone-marrow-brain in traditional Chinese medicine, we investigated the mechanism of medullary supplementation in regulating neural stem cell-derived exosomes (NSCs-Exos) to intervene in postmenopausal osteoporosis (PMOP) based on the SEMA3A-NRP1 pathway. Methods Exogenous rat neural stem cells (NSCs) were cultured in vitro and intervened with the traditional Chinese medicine Busui DAN (BSD). NSCs-Exos were extracted after 14 days of cultivation. They were divided into the group of NSCs-derived exosomes and the group of exosomes from NSCs intervened with BSD. Subsequently, transmission electron microscopy (TEM), grain size analysis (NTA), and Alix, CD9, CD63 exosome marker protein assays were performed. Animal experiments: SD rats were modeled by ovariectomy (OVX) in addition to the Control group and divided into OVX, BSD, NSCs-Exos, NSCs-Exos (BSD), NSCs-Exos+BSD, and NSCs-Exos (BSD)+BSD groups. The drug administration group was administered by gavage. The rats in the exosome group was injected with the corresponding exosome through the tail vein. The serum, kidney, femurs, and brain were taken at the end of treatment. Bone mineral density, serum ALP/TRACP, and HE staining tests were performed to analyze the effects on bone mineral density and the structural morphology of the kidney, femurs, and brain. The effects on the expressions of SEMA3A, NRP1, Runx2, and β-catenin in the kidney, femurs, and brain were analyzed by using the qRT-PCR method, ELISA method, Western blotting method, and WES method. In the same period, the DIR small animal tracking experiments in vivo were carried out to observe the targeting of NSCs-Exos. Results NSCs-Exos extracted from in vitro experiments showed round particles, which were clearly visible. CD9, CD63 and Alix were expressed, which was consistent with the characteristics of exosome. Compared to the NSCs-Exos group, the NSCs-Exos (BSD) group had a fuller morphology and higher expressions of CD9, CD63, and Alix. Animal experiments: compared to those in the control group, rats in the OVX model group had disorganized trabecular structure, significantly lower femoral bone mineral density and serum ALP expression, elevated TRACP expression, and significantly lower expressions of SEMA3A, NRP1, Runx2, and β-catenin (P<0.05), indicating that the modeling had been successful. Compared to those in the OVX model group, the femoral bone mineral density in the treatment group was significantly higher, the trabeculae number and arrangement were improved, serum ALP expression elevated, TRACP expression reduced, and SEMA3A, NRP1, Runx2, and β-catenin expressions elevated (P<0.05). The best effect was found in the NSCs-Exos (BSD)+BSD group. In the DIR experiment, it was found that NSCs-Exos expressed stronger fluorescence signal in the kidney, bone, and brain tissues, which verified its targeting. Conclusion The medullary supplement improves bone metabolism through the regulation of SEMA3A-NRP1 pathway by NSCs-Exos to prevent and control PMOP. This demonstrates the scientific connotation of the theory of kidney-bone-medullary-brain axis.
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